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The geography and ecology of the Everglades involve the complex elements affecting the natural environment throughout the southern region of the U.S. state of Florida. Before drainage, the Everglades was an interwoven mesh of marshes and prairies covering 4,000 square miles (10,000 km2). The Everglades is simultaneously a vast watershed that has historically extended from Lake Okeechobee 100 miles (160 km) south to Florida Bay (around one-third of the southern Florida peninsula), and many interconnected ecosystems within a geographic boundary. It is such a unique meeting of water, land, and climate that the use of either singular or plural to refer to the Everglades is appropriate. When Marjory Stoneman Douglas wrote her definitive description of the region in 1947, she used the metaphor "River of Grass" to explain the blending of water and plant life. Although sawgrass and sloughs are the enduring geographical icons of the Everglades, other ecosystems are just as vital, and the borders marking them are subtle or nonexistent. Pinelands and tropical hardwood hammocks are located throughout the sloughs; the trees, rooted in soil inches above the peat, marl, or water, support a variety of wildlife. The oldest and tallest trees are cypresses, whose roots are specially adapted to grow underwater for months at a time. The Big Cypress Swamp is well known for its 500-year-old cypresses, though cypress domes can appear throughout the Everglades. As the fresh water from Lake Okeechobee makes its way to Florida Bay, it meets salt water from the Gulf of Mexico; mangrove forests grow in this transitional zone, providing nursery and nesting conditions for many species of birds, fish, and invertebrates. The marine environment of Florida Bay is also considered part of the Everglades because its sea grasses and aquatic life are attracted to the constant discharge of fresh water.